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	Students have the opportunity to particpate in a research cruise on board the UNOLS research vessel<em> Atlantic Explorer </em>as part of the Modern Observational Oceanography course.</p>

Students have the opportunity to particpate in a research cruise on board the UNOLS research vessel Atlantic Explorer as part of the Modern Observational Oceanography course.

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	Students on the Coral Reef Ecology course learn commonly used techniques to study benthic community structure at several reef types.</p>

Students on the Coral Reef Ecology course learn commonly used techniques to study benthic community structure at several reef types.

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	Students take a field trip to explore Bermuda's cave systems around Castle Harbour and Blue Hole Park.</p>

Students take a field trip to explore Bermuda's cave systems around Castle Harbour and Blue Hole Park.

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	Modern Observational Oceanography students gain hands-on experience of a glider deployment five miles off shore Bermuda.</p>

Modern Observational Oceanography students gain hands-on experience of a glider deployment five miles off shore Bermuda.

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	Students on the Coral Reef Ecology course perform video surveys, fish counts and macroalgae collections. Along with associated lab work, students study the benthic community structure of reefs in Bermuda.</p>

Students on the Coral Reef Ecology course perform video surveys, fish counts and macroalgae collections. Along with associated lab work, students study the benthic community structure of reefs in Bermuda.

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	Students on the Coral Reef Ecology course separate coral tissue from the skeleton to assess physiological changes during laboratory experiments.</p>

Students on the Coral Reef Ecology course separate coral tissue from the skeleton to assess physiological changes during laboratory experiments.

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	Marine Plankton Ecology students examine taxonomy, functional diversity and the roles that plankton play in marine food webs.</p>

Marine Plankton Ecology students examine taxonomy, functional diversity and the roles that plankton play in marine food webs.

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	Summer course lectures, tutorials and workshops take place in designated lecture rooms at BIOS. </p>

Summer course lectures, tutorials and workshops take place in designated lecture rooms at BIOS. 

Each summer, BIOS offers a suite of courses for both undergraduate and graduate students that capitalize upon the expertise of our faculty and visiting scientists. These courses, listed below, provide many students the opportunity to study topics in marine science that might not be offered within the curricula of their home institutions. Each course comprises lectures, laboratory exercises, and complementary field components that build upon what is learned in the classroom.

Coral Reef Ecology

Course details TBA

Marine Larval Ecology: Responses to a Changing Ocean Environment

July 5 – July 23, 2021
Instructors:
Dr. Justin McAlister (College of the Holy Cross) and Dr. Scott Santagata (Long Island University - Post)

The larval developmental stages of many marine invertebrates are highly diverse in form, function, and life history, and effectively link benthic and pelagic ecosystems because they are the dispersive stage for many organisms, and can be particularly sensitive to environmental change. These environmental variables may include elevated ocean temperatures, increased ocean acidity, patchily distributed phytoplankton food, as well as exposure to pollutants like petroleum, heavy metals, and micro plastics. Larvae respond to stress exposure at molecular, physiological, and morphological levels. For many organisms, “normal” developmental patterns are not well known, let alone how development may vary in response to single or multiple interacting environmental stressors.

This course will examine the ecology, evolution, and development of marine invertebrate larvae, their roles as members of the meroplankton, and their responses to environmental stress at different biological scales. Students will gain hands-on experience collecting marine invertebrates from local habitats (mangrove, coral reef, pelagic open water) during boat and shore-based excursions. In the laboratory, students will learn to spawn adults, obtain and fertilize gametes, culture larvae, and conduct empirical studies of larval development under conditions of current and potential future environmental stress. Modern physiological, molecular, and microscopy-based methods will be used throughout the course. Lectures and laboratories will cover a broad range of topics and principles relevant to larval biology. More information and an example schedule can be found here.

This course is structured for upper level undergraduate and graduate students. Prerequisites include satisfactory standing in marine biology (or other relevant) courses. Snorkeling and diving opportunities will be available to those students having moderate swimming abilities and training. Course fee: $4,900 (includes tuition, room and board). Students can apply to BIOS for scholarships to assist with the course fee.

Application Instructions & Financial Aid Information

The Summer Course Application Form, along with complete instructions, can be found here. Please download the form to your computer (e.g., save it to your desktop), complete it, save it, and send it along with all supporting documents as listed on the application to education@bios.edu by May 15 in order to receive full consideration. Letters of reference should be sent directly to the BIOS education email by referees. Late applications will be considered until the courses are full. Forms filled out in browser mode (i.e., in preview) may not save appropriately. 

Partial scholarships are available for summer course fees. We do not award scholarships toward travel costs. To apply for a scholarship please fill out the appropriate section at the end of the application form and submit a statement of need along with the core application documents. All students accepted into a BIOS summer course are eligible to receive a partial scholarship from BIOS scholarship funding. Information on summer course scholarships, generally, as well as eligibility for specific scholarships, can be found on the scholarships page. Dalhousie students may apply for scholarships to attend BIOS summer courses through the Dalhousie-BIOS Experiential Learning Fund; please refer to the Dalhousie-BIOS Application for Undergraduate Scholarships for the deadline, more information and how to apply.

Please read the Course Payment Schedule & Refund Policy, as well as the note about Course Dates and Academic Credit before making your travel plans. Foreign nationals traveling to Bermuda do not require an entry visa. However, proof of residency or a valid Multi Re-Entry Visa for the U.S.A., U.K., or Canada may be required for non-U.S./U.K./Canadian citizens, depending upon your country of origin. Please review Immigration Information to check the Bermuda Department of Immigration requests.

Please note that BIOS may cancel a summer course if there is insufficient enrollment.

Monitoring the Coronavirus (COVID-19)

The health and well-being of everyone involved in BIOS’s educational programs is our top priority.

If you have questions that relate to the coronavirus disease as it concerns your plans for studying at BIOS, including application deadlines, the potential for cancellations, rescheduling, and refunding, please contact the Education Department

When making travel plans, please review updated health advisories and guidelines provided by the U.S. Department of State, the Bermuda Government, and the World Health Organization.