Coral Reef Ecology students cool off in Blue Hole.</p>

Coral Reef Ecology students cool off in Blue Hole.

	Princeton students record zonation patterns of animals and plants on the shoreline.</p>

Princeton students record zonation patterns of animals and plants on the shoreline.

	Ryan Woods and Whitney Jailley use a transect and photoquad to identify marine invertebrates on North East Breakers reef.</p>

Ryan Woods and Whitney Jailley use a transect and photoquad to identify marine invertebrates on North East Breakers reef.

	Princeton University students in the Marine Biology lab.</p>

Princeton University students in the Marine Biology lab.

	A summer student conducting a transect on the reef.</p>

A summer student conducting a transect on the reef.

	Princeton students examine Sargassum.</p>

Princeton students examine Sargassum.

Applications for 2014 summer courses are no longer being accepted.  Applications for 2015 courses will be accepted from January 2015.

Each summer, BIOS offers a suite of courses for both graduate and undergraduate students that capitalizes upon the expertise of our faculty and visiting scientists.  These courses, listed below, provide many students the opportunity to study topics in marine science that might not be offered within the curricula of their home institutions.  Each course comprises lectures, laboratory exercises, and complementary field components that build upon what is learned in the classroom. Scholarships are available to all students. 

Coral Reef Ecology: Reef Response to Environmental Change

August 4-22, 2014. Please note that students are expected to arrive the day before, and depart the day after, the course dates shown.

Open to all students. SCUBA certification is required.


Instructors: Dr. Eric Hochberg (BIOS) and Dr. Gretchen Goodbody-Gringley (BIOS)

The aim of this course is to explore the responses of coral reefs to the changing environmental conditions associated with projected global climate change scenarios. This course is offered in two components. First, a coral reef biogeochemistry section emphasizes the ecosystem processes of metabolism (primary production and respiration) and calcification of reef communities. The second section emphasizes the processes of reproduction and recruitment, response to environmental change, and resilience of reefs to human impacts. Lectures will be complemented with field and laboratory exercises. Students will gain experience in field techniques measuring metabolism and calcification, assessing recruitment recovery, monitoring bleaching, disease, invasive species and coral spawning.  Laboratory experiments will include coral fertilization, larval development and settlement, and use of a flume mesocosm to investigate effects of temperature and acidification on metabolism and calcification of different types of reef communities. A full course description can be found here, and a draft schedule can be found hereCourse fee in 2014: $4,900 (tuition, room and board). Students can apply to BIOS for scholarships to assist with the course fee.

Tropical Marine Ecology and Conservation

Course not offered in 2014.

Open to all students. 

Instructors: n/a

Students in the Tropical Marine Ecology and Conservation course will study the features, functions and interactions of tropical marine ecosystems and the challenges facing their conservation. Through lectures, discussions and integrated field work and laboratory exercises, students will examine the characteristics, biodiversity, biogeography and ecology of coral reef, seagrass, mangrove and other coastal habitats. Human exploitation and global climate change impacts will be examined along with conservation biology and resource management strategies. The course is designed to provide practice in a variety of marine ecological field sampling and laboratory techniques. BIOS has quick and easy access to a diverse array of tropical marine habitats and well-equipped labs providing an optimal environment for experiential learning. A full course description can be found hereCourse fee in 2014: $4,900 (tuition, room and board). Students can apply to BIOS for scholarships to assist with the course fee.

Marine Biology (EEB/ENV 312, held at BIOS)

June 1-28, 2014 

Open to Princeton University students of Ecology and Evolutionary Biology and those attending Princeton Environmental Institute (PEI) programs.


Instructors: Dr. Samantha dePutron (BIOS) and Dr. James Gould (Princeton University)

A field course for Princeton undergraduate students. This intensive four-week course in June will cover all aspects of marine biology, with a special focus on marine ecology and coral reefs. Other habitats include open ocean, seagrass beds, coastal zone, rocky shore, marshes and marine ponds, and mangroves. The topics include the ecology, physiology, and behavior of the resident organisms, which will range from plankton to whales, with special attention to corals, marine plants, and reef fish. For complete course description and details see here, and please visit the Department of Ecology and Evolutionary Biology's Marine Biology Summer website. Tentative course schedule for 2014.

Students have commented that they “loved the field work and the opportunity to see the whole island. It was extremely interesting to work alongside other researchers and amazing faculty.” One student said, “I learned more in a four-week course than I do in most semester-long classes.”

Microbial Oceanography: The Biogeochemistry, Ecology and Genomics of Oceanic Microbial Ecosystems

Course not offered in 2014

Open to all undergraduate and graduate students. SCUBA certification is not required.


Instructors: Dr. Craig Carlson, University of California, Santa Barbara; Dr. Stephen Giovannoni, Oregon State University; Dr. John Heidelberg, University of Southern California; Dr. Craig E. Nelson, University of California, Santa Barbara

The course is offered with two major components: Microbial Molecular Ecology and Microbial Genetics.  The Marine Microbial Molecular Ecology course will  cover traditional microbiological topics such as physiology, microbial diversity and growth within the context of biogeochemical processes.  Lectures will focus on how biological processes and the ecological structure within the water column control the cycling of important elements in the ocean such as carbon, nitrogen, and phosphorus.  A series of field trips and laboratory experiments will be conducted to investigate microbial processes in the open ocean and in the coral reef dominated waters of Bermuda. The Marine Microbial Genomics course will introduce the types of data that can be obtained from modern genomic methods (e.g., survey sequencing and "meta-" or community genomics of entire ecosystems) and the computational tools useful in the analysis of sequence data.

Nicholas Huynh, a 2013 course attendee said, “This intensive course taught by some of the brightest minds in microbial oceanography has been greatly beneficial to my pursuit of a career in this field and has been influential in the way that I think. It has been an extraordinary opportunity to work alongside an international group of like-minded individuals. I am leaving Bermuda with a stronger sense of how to approach and conduct microbial biogeochemical research as a team member.”

Application Instructions & Financial Aid Information

The 2014 Summer Course Application Form, along with complete instructions, can be found here. Please download the form to your computer (e.g., save it to your desktop), complete it, save it, and send it along with all supporting documents listed on the application to education@bios.edu by April 30, 2014 in order to receive full consideration. Late applications will be considered until the courses are full. Forms filled out in browser mode (i.e., in preview) may not save appropriately.  

Limited partial scholarships are available for summer course tuition; we do not award scholarships toward travel. To apply for a scholarship please fill out the appropriate section at the end of the application form and submit a statement of need along with the core application documents. For information about scholarships to attend summer courses at BIOS, please visit the Undergraduate Scholarships page. Dalhousie student may apply for scholarships to attend BIOS summer courses through the Dalhousie-BIOS Experiential Learning Fund; please refer to the Dalhousie-BIOS Application for Undergraduate Scholarships for more information and how to apply.

Please read the Course Payment Schedule & Refund Policy, as well as the note about Course Dates and Academic Credit before making your travel plans. Foreign nationals traveling to Bermuda do not require an entry visa. However, proof of residency or a valid Multi Re-Entry Visa for the USA, UK, or Canada may be required for non-US/UK/Canadian citizens, depending upon your country of origin. Please review Immigration Information to check the Bermuda Department of Immigration requests.